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                            PHARAOH ANT


Introduction

The ant, Monomorium pharaonis (Linnaeus), is commonly known as the Pharaoh ant. The name possibly arises from the mistaken tradition that it was one of the plagues of ancient Egypt (Peacock et al. 1950). This ant is distributed worldwide, is one of the more common household ants, and carries the dubious distinction of being the most difficult household ant to control.


In some of the older literature this species was commonly referred to as the "Pharaoh's ant." This is now incorrect, if it ever was, as the correct common name is "Pharaoh ant," as determined by the Committee on Common Names of Insects of the Entomological Society of America (Bosik 1997). It is only mentioned here as the junior author still sees "Pharaoh's ant" on the Web and in industry publications.

Distribution

Monomorium pharaonis (Linnaeus) has been carried by commerce to all inhabited regions of the earth (Wheeler 1910). This ant, which is probably a native of Africa, does not nest outdoors except in southern latitudes and has been able to adapt to field conditions in southern Florida (Creighton 1950). In colder climates, it has become established in heated buildings.

Description

The workers of Monomorium pharaonis (L.) while monomorphic (same size), do vary slightly in length and are approximately 1.5 to 2 mm long (Haack and Granovsky 1990). The antennae have 12 segments with each segment of the 3-segmented antennal clubs increasing in size toward the apex of the club (Smith and Whitman 1992). The eye is comparatively small, with approximately six to eight ommatidia across the greatest diameter. The prothorax has subangular shoulders, and the thorax has a well- defined mesoepinotal impression. Erect hairs are sparse on the body, and body pubescence is sparse and closely appresssed. The head, thorax, petiole and postpetiole (the petiole, or the petiole and postpetiole, in ants is also called the pedicel) are densely (but weakly) punctulate, dull, or subopaque. The clypeus, gaster, and mandibles are shiny. The body color ranges from yellowish or light brown to red (Smith 1965), with the abdomen often darker to blackish (Smith and Whitman 1992). A stinger is present but is rarely exserted (Haack and Granovsky 1990).

Biology and Nesting Habits

The Pharaoh ant colony consists of queens, males, workers, and immature stages (eggs, larvae, pre- pupae, and pupae). Nesting occurs in inaccessible warm (80 to 86°F), humid (80%) areas near sources of food and/or water, such as in wall voids. The size of the colony tends to be large but can vary from a few dozen to several thousand or even several hundred thousand individuals. Approximately 38 days are required for development of workers from egg to adult.

Mating takes place in the nest, and no swarms are known to occur. Males and queens usually take 42 days to develop from egg to adult. The males are the same size as the workers (2 mm), are black in color and have straight, not elbowed, antennae. Males are not often found in the colony. The queens are about 4 mm long and are slightly darker than the workers (Smith and Whitman 1992). Queens can produce 400 or more eggs in batches of 10 to 12 (Peacock et al. 1950). Queens can live four to 12 months, while males die within three to five weeks after mating (Smith and Whitman 1992).

Part of the success and persistence of this ant undoubtably relates to the budding or splitting habits of the colonies. Numerous daughter colonies are produced from the mother colony when a queen and a few workers break off and establish a new colony. Even in the absence of a queen, workers can develop a queen from the brood which is transported from the mother country. In large colonies there may be as many as several hundred reproductive females (Smith and Whitman 1992).

Economic Importance

The Pharaoh ant is a major indoor pest in the United States. The ant has the ability to survive most conventional household pest control treatments and to establish colonies throughout a building. More than just the food it consumes or spoils, this ant is considered a serious pest simply due to its ability for "getting into things." Pharaoh ants are reported to have even penetrated the security of recombinant DNA laboratories (Haack and Granovsky 1990).

In some areas, this ant has become a major pest of residences, commercial bakeries, factories, office buildings, apartments, and hospitals or other areas where food is handled. Infestations in hospitals have become a chronic problem in Europe (Erodes et al. 1977) and the United States. In Texas, Wilson and Booth (1981) reported an extensive infestation throughout a seven-floor medical center. In ant- infested hospitals, burn victims and newborns are subjected to increased risk because the Pharaoh ant can transmit over a dozen pathogenic pathogens such as Salmonella spp., Staphylococcus spp., and Streptococcus spp. (Beatson 1977, Haack and Granovsky 1990, Smith and Whitman 1992). Pharaoh ants have been observed seeking moisture from the mouths of sleeping infants and from in-use IV bottles (Smith and Whitman 1992).

This ant infests almost all areas of a building where food is available and infests many areas where food is not commonly found. Pharaoh ants have a wide preference in the types of food consumed. In infested areas, if sweet, fatty, or oily foods are left uncovered for only a short period of time, one can likely find a trail of Pharaoh ants to the food. As a consequence, they cause much food to be discarded due to contamination. Owners have been known to consider selling their homes because of the ravages of this pest (Smith 1965).


Survey and Detection

Workers of the Pharaoh ant can often be observed on their feeding trails, often using wiring or hot water pipes to travel through walls and between floors. Once a worker has located a food source, it lays a chemical trail from the food to the nest (Haack and Granovsky 1990). These ants are attracted to sweet and fatty foods, which may be used to determine their presence. Pharaoh ants will nest in the oddest places, such as between sheets of stationary, layers of bed linen and clothes, in appliances, or even piles of trash (Ebeling 1978).

Pharaoh ants may be confused with thief ants, bigheaded ants, firs ants and several other species of small pale ants. However, thief ants have just 10 segments in their antennae with only a 2-segmented club. Bigheaded and fire ants have a pair of spines on the thorax, while other small pale ants have only a one segment on the pedical (smith and Whitman 1992).


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